@import url('https://fonts.googleapis.com/css2?family=Cormorant+Garamond:ital,wght@0,300;0,400;0,500;1,300;1,400;1,500&family=Inter:wght@400;500;600&display=swap');

/* ============================================================
   Kaplan Lab — Stylesheet v2
   University of Virginia, Department of Biology
   ============================================================ */

:root {
  --dark:        #08152A;
  --navy:        #0D2B55;
  --navy-mid:    #1E4080;
  --amber:       #C8870A;
  --amber-pale:  #FEF3C7;
  --white:       #FFFFFF;
  --bg:          #FAFBFD;
  --off-white:   #FAFAF7;
  --surface:     #F1F4FA;
  --text:        #0F172A;
  --text-mid:    #374151;
  --muted:       #6B7280;
  --border:      #E2E8F2;
  --border-mid:  #CDD6E8;

  --font-display: 'Cormorant Garamond', Georgia, serif;
  --font-serif:   'Cormorant Garamond', Georgia, serif;
  --font-sans:    'Inter', system-ui, -apple-system, sans-serif;

  --max-w:  1100px;
  --nav-h:  72px;

  --sp-xs:  8px;
  --sp-sm:  16px;
  --sp-md:  28px;
  --sp-lg:  48px;
  --sp-xl:  80px;
  --sp-2xl: 120px;

  --r-sm: 6px;
  --r-md: 14px;
  --r-lg: 20px;
  --r-xl: 28px;

  --shadow-sm: 0 1px 4px rgba(8,21,42,.06);
  --shadow-md: 0 4px 24px rgba(8,21,42,.09);
  --shadow-lg: 0 12px 48px rgba(8,21,42,.13);
}

/* ---- Reset ---- */
*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }
html { scroll-behavior: smooth; -webkit-text-size-adjust: 100%; }
body {
  font-family: var(--font-sans);
  font-size: 1rem;
  line-height: 1.65;
  color: var(--text);
  background: var(--off-white);
}
img, video { max-width: 100%; height: auto; display: block; }
a { color: var(--navy); text-decoration: none; transition: color .2s; }
a:hover { color: var(--amber); }
button { font-family: var(--font-sans); }

/* ---- Typography ---- */
h1, h2, h3 {
  font-family: var(--font-display);
  font-weight: 400;
  line-height: 1.12;
}
h4, h5 { font-family: var(--font-sans); font-weight: 600; line-height: 1.3; }

h1 { font-size: clamp(3.2rem, 7.5vw, 6.5rem); }
h2 { font-size: clamp(2.4rem, 5vw, 3.8rem); }
h3 { font-size: clamp(1.4rem, 2.5vw, 2rem); }
h4 { font-size: 1rem; }
h5 { font-size: .75rem; text-transform: uppercase; letter-spacing: .12em; }

p { max-width: 65ch; }

.lead {
  font-size: 1.12rem;
  line-height: 1.75;
  color: var(--muted);
}

.intro-statement {
  font-family: var(--font-display);
  font-size: clamp(1.5rem, 3vw, 2.4rem);
  font-style: italic;
  font-weight: 300;
  color: var(--text-mid);
  line-height: 1.4;
  text-align: center;
  max-width: 760px;
  margin: 0 auto;
}

/* Label */
.label, .section-label {
  display: block;
  font-size: 1rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: .18em;
  color: var(--navy-mid);
  margin-bottom: var(--sp-sm);
}

/* ---- Layout ---- */
.container {
  width: 100%;
  max-width: var(--max-w);
  margin: 0 auto;
  padding: 0 var(--sp-lg);
}

.section          { padding: var(--sp-2xl) var(--sp-lg); }
.section--alt     { background: var(--surface); }
#team.section { padding-top: calc(var(--sp-xl) * 0.8); padding-bottom: var(--sp-xl); }
#team { scroll-margin-top: calc(var(--nav-h) + 12px); }
.section--dark    { background: var(--dark); color: var(--white); }
.section--navy    { background: var(--navy); color: var(--white); }
.section--border  { border-top: 1px solid var(--border); }

.section-intro { max-width: 640px; margin-bottom: var(--sp-xl); }
.section-intro h2 { margin-top: var(--sp-xs); }
.section-intro .lead { margin-top: var(--sp-md); }

.flex-between {
  display: flex;
  justify-content: space-between;
  align-items: baseline;
  flex-wrap: wrap;
  gap: var(--sp-md);
}

/* ---- Navigation ---- */
.site-nav {
  position: fixed;
  top: 0; left: 0; right: 0;
  height: var(--nav-h);
  z-index: 200;
  background: rgba(255,255,255,.97);
  backdrop-filter: blur(18px);
  -webkit-backdrop-filter: blur(18px);
  border-bottom: 1px solid var(--border);
  transition: box-shadow .4s ease;
}
.site-nav.scrolled {
  box-shadow: 0 1px 24px rgba(8,21,42,.07);
}

.site-nav__inner {
  display: flex;
  align-items: center;
  justify-content: space-between;
  height: 100%;
  max-width: var(--max-w);
  margin: 0 auto;
  padding: 0 var(--sp-lg);
}

.site-nav__logo {
  display: flex;
  flex-direction: column;
  gap: 2px;
  text-decoration: none;
  opacity: 0;
  pointer-events: none;
  transform: translateY(-6px);
  transition: opacity .2s ease, transform .2s ease;
}
.site-nav.scrolled .site-nav__logo {
  opacity: 1;
  pointer-events: auto;
  transform: translateY(0);
}
.site-nav__logo-main {
  font-family: var(--font-display);
  font-size: 1.25rem;
  color: var(--text);
  line-height: 1;
  letter-spacing: .01em;
}
.site-nav__logo-sub {
  font-size: .58rem;
  color: var(--muted);
  letter-spacing: .12em;
  text-transform: uppercase;
}

.site-nav__links { display: flex; list-style: none; gap: clamp(16px, 2.8vw, 44px); }
.site-nav__links a {
  font-size: clamp(.72rem, .9vw, .82rem);
  font-weight: 500;
  color: var(--muted);
  letter-spacing: .06em;
  text-transform: uppercase;
  padding-bottom: 3px;
  border-bottom: 1.5px solid transparent;
  transition: color .2s, border-color .2s;
  white-space: nowrap;
}
.site-nav__links a:hover { color: var(--text); }
.site-nav__links a.active {
  color: var(--navy);
  border-bottom-color: var(--navy);
  font-weight: 600;
}

.site-nav__hamburger { display: none; background: none; border: none; cursor: pointer; padding: 6px; }
.site-nav__hamburger span {
  display: block; width: 22px; height: 1.5px;
  background: var(--navy);
  margin: 5px 0;
  transition: transform .3s, opacity .3s, background .4s;
  border-radius: 2px;
}
.site-nav.scrolled .site-nav__hamburger span { background: var(--text-mid); }

/* ---- Hero (home) ---- */
.hero {
  position: relative;
  height: 100vh;
  min-height: 580px;
  display: flex;
  flex-direction: column;
  background: var(--bg);
  overflow: hidden;
}


.hero__content {
  position: relative; z-index: 2;
  width: 100%;
  padding: calc(var(--nav-h) + 40px) var(--sp-lg) var(--sp-lg);
}

/* ---- Hero image gallery ---- */
.hero__gallery {
  position: relative;
  width: 100%;
  flex: 1;
  min-height: 0;
  z-index: 1;
  display: flex;
  align-items: center;
  justify-content: center;
}
.hero__gallery-slide {
  position: absolute;
  inset: 0;
  opacity: 0;
  pointer-events: none;
  display: flex;
  align-items: center;
  justify-content: center;
  background: var(--bg);
  animation: hero-crossfade 10.5s ease-in-out infinite;
}
.hero__gallery-slide:nth-child(1) { animation-delay: 0s; }
.hero__gallery-slide:nth-child(2) { animation-delay: -7.0s; }
.hero__gallery-slide:nth-child(3) { animation-delay: -3.5s; }
.hero__gallery-slide img {
  width: 100%;
  height: 100%;
  object-fit: contain;
  display: block;
  max-width: 100%;
}
@keyframes hero-crossfade {
  0%, 29%  { opacity: 1; }
  33%, 95% { opacity: 0; }
  100%     { opacity: 1; }
}

.hero__gallery-ph {
  width: 100%; height: 100%;
  display: flex; align-items: center; justify-content: center;
  font-size: .72rem; font-weight: 500;
  letter-spacing: .14em; text-transform: uppercase;
  color: rgba(255,255,255,.2);
  border-top: 1px solid rgba(255,255,255,.06);
}

.hero__eyebrow {
  display: inline-flex;
  align-items: center;
  gap: 14px;
  font-size: .88rem;
  font-weight: 600;
  color: var(--amber);
  text-transform: uppercase;
  letter-spacing: .16em;
  margin-bottom: var(--sp-md);
}
.hero__eyebrow::before {
  content: '';
  display: block;
  width: 36px; height: 1px;
  background: var(--amber);
}

.hero__lab-title {
  font-family: var(--font-display);
  font-size: clamp(3rem, 6vw, 4.4rem);
  color: var(--navy);
  margin: 0 0 .45rem;
  line-height: 1.02;
  letter-spacing: .01em;
}

.hero__headline {
  color: var(--dark);
  max-width: 100%;
  margin-bottom: 0;
  font-weight: 400;
  letter-spacing: -.02em;
  font-size: clamp(2rem, 3.5vw, 3.5rem);
  line-height: 1.22;
}
.hero__headline em {
  font-style: italic;
  font-weight: 400;
  color: var(--amber);
}
.hero__sep {
  color: var(--border-mid);
  font-style: normal;
  margin: 0 .15em;
}

.hero__body {
  font-size: 1.05rem;
  color: rgba(255,255,255,.58);
  max-width: 480px;
  margin-bottom: var(--sp-xl);
  line-height: 1.8;
}
.hero__body p { max-width: none; }

.hero__cta { display: flex; gap: 14px; flex-wrap: wrap; }

.hero__meta {
  position: absolute;
  bottom: 40px; left: var(--sp-lg);
  right: var(--sp-lg);
  max-width: var(--max-w);
  margin: 0 auto;
  display: flex;
  justify-content: space-between;
  align-items: flex-end;
  z-index: 2;
}
.hero__affil {
  font-size: .72rem;
  color: rgba(255,255,255,.28);
  letter-spacing: .08em;
}
.hero__scroll {
  display: flex;
  flex-direction: column;
  align-items: center;
  gap: 8px;
  color: rgba(255,255,255,.28);
  font-size: .62rem;
  letter-spacing: .18em;
  text-transform: uppercase;
}
.hero__scroll-line {
  width: 1px; height: 40px;
  background: linear-gradient(to bottom, transparent, rgba(255,255,255,.28));
  animation: scrollPulse 2.4s ease-in-out infinite;
}
@keyframes scrollPulse {
  0%,100% { transform: scaleY(1); opacity: .4; }
  50%      { transform: scaleY(1.25); opacity: 1; }
}

/* ---- Page hero (interior pages) ---- */
.page-hero {
  background: var(--dark);
  padding: calc(var(--nav-h) + var(--sp-xl)) var(--sp-lg) var(--sp-xl);
}
.page-hero__inner { max-width: var(--max-w); margin: 0 auto; }
.page-hero h1 { color: var(--white); margin: var(--sp-xs) 0 var(--sp-md); letter-spacing: -.02em; }
.page-hero .lead { color: rgba(255,255,255,.55); max-width: 62ch; }

/* ---- Buttons ---- */
.btn {
  display: inline-flex;
  align-items: center;
  gap: 8px;
  padding: 13px 26px;
  border-radius: var(--r-sm);
  font-size: .86rem;
  font-weight: 500;
  font-family: var(--font-sans);
  cursor: pointer;
  border: none;
  text-decoration: none;
  transition: all .22s ease;
  line-height: 1;
  letter-spacing: .02em;
}
.btn--primary {
  background: var(--amber);
  color: var(--dark);
}
.btn--primary:hover {
  background: #D9920C;
  color: var(--dark);
  transform: translateY(-1px);
  box-shadow: 0 6px 22px rgba(200,135,10,.32);
}
.btn--outline {
  background: transparent;
  color: var(--white);
  border: 1px solid rgba(255,255,255,.3);
}
.btn--outline:hover {
  background: rgba(255,255,255,.07);
  color: var(--white);
  border-color: rgba(255,255,255,.6);
}
.btn--ghost {
  background: transparent;
  color: var(--text-mid);
  border: 1px solid var(--border);
  font-size: .82rem;
}
.btn--ghost:hover {
  background: var(--surface);
  color: var(--text);
  border-color: var(--border-mid);
}
.btn--navy {
  background: var(--navy);
  color: var(--white);
}
.btn--navy:hover {
  background: var(--navy-mid);
  color: var(--white);
  transform: translateY(-1px);
}

/* ---- Research list (home) ---- */
.research-list {
  border-top: 1px solid var(--border);
  margin-top: var(--sp-xl);
}
.research-list-item {
  display: grid;
  grid-template-columns: 68px 1fr 32px;
  gap: var(--sp-lg);
  padding: var(--sp-lg) var(--sp-sm);
  border-bottom: 1px solid var(--border);
  color: var(--text);
  text-decoration: none;
  align-items: center;
  transition: background .2s ease, padding-left .28s ease;
}
.research-list-item:hover {
  background: var(--surface);
  padding-left: var(--sp-md);
}
.research-list-item__num {
  font-family: var(--font-display);
  font-size: 2.6rem;
  font-weight: 300;
  color: var(--amber);
  line-height: 1;
  opacity: .55;
}
.research-list-item h3 {
  font-size: clamp(1.5rem, 2.5vw, 2rem);
  margin-bottom: 8px;
  color: var(--text);
  font-weight: 400;
  letter-spacing: -.01em;
}
.research-list-item p {
  font-size: .9rem;
  color: var(--muted);
  line-height: 1.62;
  max-width: 58ch;
}
.research-list-item__arrow {
  font-size: 1rem;
  color: var(--border-mid);
  transition: transform .25s, color .25s;
  justify-self: end;
}
.research-list-item:hover .research-list-item__arrow {
  transform: translateX(6px);
  color: var(--amber);
}

/* ---- Research hero block (item 01 — full-bleed 50/50) ---- */
.research-hero-block {
  display: grid;
  grid-template-columns: 1fr 1fr;
  min-height: 65vh;
  border-top: 1px solid var(--border);
  border-bottom: 1px solid var(--border);
}
.research-hero-block__text {
  padding: clamp(48px, 6vw, 88px) clamp(48px, 5vw, 80px);
  display: flex;
  flex-direction: column;
  justify-content: center;
  gap: var(--sp-md);
  border-right: 1px solid var(--border);
}
.research-hero-block__num {
  font-family: var(--font-sans);
  font-size: .85rem;
  font-weight: 700;
  letter-spacing: .12em;
  text-transform: uppercase;
  color: var(--navy-mid);
}
.research-hero-block__text h3 {
  font-family: var(--font-serif);
  font-size: clamp(1.9rem, 2.8vw, 2.9rem);
  font-weight: 400;
  line-height: 1.18;
  color: var(--text);
  letter-spacing: -.01em;
  margin: 0;
}
.research-hero-block__text p {
  font-size: clamp(.95rem, 1.15vw, 1.08rem);
  line-height: 1.85;
  color: var(--muted);
  max-width: 50ch;
  margin: 0;
}
.research-hero-block__video {
  background: var(--bg);
  overflow: hidden;
  border-bottom: none;
}
.research-hero-block:first-of-type .research-hero-block__video {
  position: relative;
}
.research-hero-block:first-of-type .research-hero-block__video::after {
  content: '';
  position: absolute;
  left: 0;
  right: 0;
  bottom: 0;
  height: 8px;
  background: var(--bg);
  z-index: 4;
  pointer-events: none;
}
.research-hero-block:first-of-type .research-hero-block__video::before {
  content: '';
  position: absolute;
  left: 0;
  right: 0;
  bottom: 1.2%;
  height: 3px;
  background: var(--bg);
  z-index: 4;
  pointer-events: none;
}
.research-hero-block__video video {
  width: 100%;
  height: 100%;
  object-fit: contain;
  position: relative;
  z-index: 1;
  transform: scale(1.008);
  transform-origin: center;
  display: block;
}
.research-hero-block__video img {
  width: 100%;
  height: 100%;
  object-fit: contain;
  display: block;
}
@media (max-width: 900px) {
  .research-hero-block { grid-template-columns: 1fr; min-height: auto; }
  .research-hero-block__text { border-right: none; border-bottom: 1px solid var(--border); padding: var(--sp-xl) var(--sp-lg); }
  .research-hero-block__video { aspect-ratio: 16/9; min-height: 0; }
}

/* ---- Nature cover art ---- */
.cover-art {
  flex-shrink: 0;
  width: 200px;
}
.cover-art img {
  width: 100%;
  display: block;
  border-radius: var(--r-md);
  box-shadow: var(--shadow-lg);
}
.cover-art-placeholder {
  width: 200px;
  flex-shrink: 0;
  background: var(--surface);
  border: 1.5px dashed var(--border-mid);
  border-radius: var(--r-md);
  aspect-ratio: 3/4;
  display: flex;
  flex-direction: column;
  align-items: center;
  justify-content: center;
  gap: 8px;
  padding: var(--sp-md);
  text-align: center;
  color: var(--muted);
  font-size: .78rem;
  font-style: italic;
}
.pub-hero {
  display: flex;
  gap: var(--sp-lg);
  align-items: stretch;
  margin-bottom: var(--sp-lg);
  min-height: 288px;
}
.cover-art-placeholder {
  width: min(320px, 34%);
  flex-shrink: 0;
  aspect-ratio: 3 / 4;
  height: auto;
  border-radius: var(--r-xl);
}
.cover-art-placeholder img {
  width: 100%;
  height: 100%;
  object-fit: cover;
  display: block;
}
@media (max-width: 640px) {
  .pub-hero { flex-direction: column; }
  .cover-art, .cover-art-placeholder { width: 100%; }
}

/* ---- Featured paper ---- */
.fp {
  display: grid;
  grid-template-columns: 130px 1fr;
  background: var(--dark);
  border-radius: var(--r-xl);
  overflow: hidden;
  margin-top: var(--sp-lg);
  box-shadow: var(--shadow-lg);
  min-height: 100%;
  align-content: center;
}
.fp__left {
  background: rgba(255,255,255,.025);
  border-right: 1px solid rgba(255,255,255,.07);
  padding: var(--sp-xl) var(--sp-lg);
  display: flex;
  flex-direction: column;
  align-items: center;
  justify-content: center;
  gap: 8px;
}
.fp__journal {
  font-family: var(--font-display);
  font-size: 2rem;
  color: #E9B43A;
  line-height: 1;
  letter-spacing: -.01em;
}
.fp__year {
  font-size: .68rem;
  letter-spacing: .14em;
  text-transform: uppercase;
  color: rgba(255,255,255,.28);
}
.fp__right { padding: var(--sp-xl); display: flex; flex-direction: column; justify-content: center; }
.fp__title {
  font-family: var(--font-display);
  font-size: clamp(1.5rem, 2.5vw, 2.2rem);
  color: var(--white);
  line-height: 1.28;
  margin-bottom: 12px;
  font-weight: 400;
  letter-spacing: -.01em;
}
.fp__authors {
  font-size: .82rem;
  color: rgba(255,255,255,.4);
  margin-bottom: var(--sp-md);
  letter-spacing: .01em;
}
.fp__excerpt {
  display: none;
}
.fp__excerpt p { max-width: none; }
.fp__link {
  font-size: .84rem;
  font-weight: 600;
  color: #E9B43A;
  letter-spacing: .02em;
  display: inline-flex;
  align-items: center;
  gap: 6px;
  transition: color .2s, gap .2s;
}
.fp__link:hover { color: #F2C75E; gap: 10px; }

/* ---- News list (home page) ---- */
.news-list { border-top: 1px solid var(--border); }
.news-list-item {
  display: grid;
  grid-template-columns: 88px 108px 1fr 20px;
  gap: var(--sp-md);
  padding: 20px var(--sp-sm);
  border-bottom: 1px solid var(--border);
  align-items: center;
  color: var(--text);
  text-decoration: none;
  transition: background .18s, padding-left .24s;
}
.news-list-item:hover {
  background: var(--off-white);
  padding-left: var(--sp-md);
}
.news-list-item__date {
  font-size: .75rem;
  font-weight: 600;
  color: var(--muted);
  text-transform: uppercase;
  letter-spacing: .07em;
  white-space: nowrap;
}
.news-list-item__title {
  font-size: .95rem;
  color: var(--text);
  line-height: 1.4;
}
.news-list-item__title em { font-style: italic; }
.news-list-item__arrow {
  color: var(--border-mid);
  font-size: .85rem;
  transition: transform .2s, color .2s;
}
.news-list-item:hover .news-list-item__arrow {
  transform: translateX(4px);
  color: var(--amber);
}

/* ---- Tags ---- */
.tag {
  display: inline-block;
  font-size: .67rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: .07em;
  padding: 4px 11px;
  border-radius: 100px;
  white-space: nowrap;
}
.tag--lab         { background: #FEF9EC; color: #78350F; border: 1px solid #FDE68A; }
.tag--publication { background: #EEF2FF; color: #3730A3; border: 1px solid #C7D2FE; }
.tag--talk        { background: #ECFDF5; color: #065F46; border: 1px solid #A7F3D0; }
.tag--award       { background: #F5F3FF; color: #5B21B6; border: 1px solid #DDD6FE; }
.tag--preprint    { background: #FFFBEB; color: #92400E; border: 1px solid #FDE68A; }

/* ---- Join CTA section ---- */
.cta-section { padding: var(--sp-2xl) var(--sp-lg); }
.cta-section__inner { max-width: 680px; margin: 0 auto; text-align: center; }
.cta-section h2 { color: var(--white); margin: var(--sp-xs) 0 var(--sp-md); }
.cta-section .lead { color: rgba(255,255,255,.55); max-width: 48ch; margin: 0 auto var(--sp-lg); }

/* ---- Callout ---- */
.callout {
  background: var(--amber-pale);
  border-left: 3px solid var(--amber);
  border-radius: 0 var(--r-sm) var(--r-sm) 0;
  padding: var(--sp-md) var(--sp-lg);
  margin: var(--sp-lg) 0;
}
.callout p {
  font-family: var(--font-display);
  font-size: 1.25rem;
  font-style: italic;
  color: var(--dark);
  line-height: 1.62;
  max-width: 70ch;
}

/* ---- Research areas (research.html) ---- */
.research-area {
  display: grid;
  grid-template-columns: 1fr 1fr;
  gap: var(--sp-xl);
  align-items: start;
  padding: var(--sp-2xl) 0;
  border-bottom: 1px solid var(--border);
}
.research-area:last-child { border-bottom: none; }
.research-area--flip { direction: rtl; }
.research-area--flip > * { direction: ltr; }

.research-area__num {
  font-family: var(--font-display);
  font-size: 7rem;
  line-height: 1;
  color: var(--navy-mid);
  opacity: .15;
  margin-bottom: var(--sp-sm);
  font-weight: 300;
}
.research-area__tag {
  font-size: .68rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: .16em;
  color: var(--navy-mid);
  display: block;
  margin-bottom: 10px;
}
.research-area h3 { margin-bottom: var(--sp-sm); letter-spacing: -.01em; }
.research-area__body { font-size: .95rem; color: var(--muted); line-height: 1.8; margin-bottom: var(--sp-md); }
.research-area__body p { max-width: none; margin-bottom: 14px; }
.research-area__qs { list-style: none; margin-top: var(--sp-md); }
.research-area__qs li {
  display: flex; gap: 10px;
  font-size: .88rem; color: var(--muted);
  margin-bottom: 8px; line-height: 1.5;
}
.research-area__qs li::before { content: '—'; color: var(--amber); flex-shrink: 0; font-weight: 700; }

.research-visual {
  background: var(--surface);
  border: 1px solid var(--border);
  border-radius: var(--r-lg);
  overflow: hidden;
  aspect-ratio: 4/3;
  display: flex;
  align-items: center;
  justify-content: center;
  position: relative;
}
.research-visual-placeholder {
  text-align: center;
  padding: var(--sp-xl);
}
.research-visual-placeholder svg {
  opacity: .2;
  margin: 0 auto var(--sp-sm);
}
.research-visual-placeholder p {
  font-size: .78rem;
  font-style: italic;
  color: var(--border-mid);
  max-width: 26ch;
  margin: 0 auto;
  line-height: 1.5;
}

/* ---- Approach cards ---- */
.approach-grid {
  display: grid;
  grid-template-columns: repeat(4, 1fr);
  gap: var(--sp-md);
}
.approach-card {
  background: var(--white);
  border: 1px solid var(--border);
  border-radius: var(--r-md);
  padding: var(--sp-lg);
  transition: box-shadow .22s, transform .22s;
}
.approach-card:hover { box-shadow: var(--shadow-md); transform: translateY(-3px); }
.approach-card__icon {
  width: 44px; height: 44px;
  border-radius: var(--r-sm);
  background: var(--amber-pale);
  display: flex; align-items: center; justify-content: center;
  margin-bottom: var(--sp-sm);
  font-size: 1.2rem;
}
.approach-card h4 { margin-bottom: 8px; font-size: .97rem; }
.approach-card p  { font-size: .87rem; color: var(--muted); line-height: 1.65; }



.pub-year-group { margin-bottom: 10px; }
.pub-year-group__label {
  font-family: var(--font-display);
  font-size: 2.1rem;
  color: var(--navy);
  border-bottom: 2px solid var(--navy);
  display: inline-block;
  padding-bottom: 3px;
  margin-bottom: 8px;
  font-weight: 400;
}

.pub-item {
  display: flex;
  gap: var(--sp-lg);
  padding: 8px 0;
  border-bottom: 1px solid var(--border);
}
.pub-item:last-child { border-bottom: none; }
.pub-item__meta { min-width: 52px; }
.pub-item__badge {
  display: inline-flex;
  padding: 3px 10px;
  border-radius: 100px;
  font-size: .67rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: .05em;
  white-space: nowrap;
}
.badge--nature   { background: #000;    color: #fff; }
.badge--cell     { background: #003566; color: #fff; }
.badge--neuron   { background: #1D4ED8; color: #fff; }
.badge--phrv     { background: #6D28D9; color: #fff; }
.badge--currop   { background: #047857; color: #fff; }
.badge--elife    { background: #0891B2; color: #fff; }
.badge--plos     { background: #B45309; color: #fff; }
.badge--crm      { background: #DC2626; color: #fff; }
.badge--biorxiv  { background: #92400E; color: #fff; }
.badge--ptrsl    { background: #4B5563; color: #fff; }

.pub-item__content { flex: 1; }
.pub-item__journal-line { display: flex; align-items: center; gap: 8px; margin-bottom: 7px; }
.pub-item__journal-info { font-size: .8rem; color: var(--muted); }
.pub-item__title { font-size: .97rem; font-weight: 600; line-height: 1.4; margin-bottom: 4px; }
.pub-item__title a { color: inherit; }
.pub-item__title a:hover { color: var(--navy-mid); }
.pub-item__authors { font-size: .85rem; color: var(--muted); line-height: 1.5; overflow-wrap: normal; }
.pub-item__authors strong { color: var(--navy); font-weight: 600; text-decoration: underline; }
.pub-item__authors sup,
.fp__authors sup {
  font-size: .72em;
  line-height: 0;
  vertical-align: super;
}
.eq-contrib {
  white-space: nowrap;
}
.pub-item__links { display: flex; gap: 8px; margin-top: 6px; flex-wrap: wrap; }
.pub-link {
  font-size: .73rem;
  font-weight: 600;
  color: var(--navy);
  background: var(--surface);
  padding: 3px 10px;
  border-radius: var(--r-sm);
  transition: all .2s;
}
.pub-link:hover { background: var(--navy); color: var(--white); }

/* ---- Member grid ---- */
.member-grid {
  display: grid;
  grid-template-columns: repeat(auto-fill, minmax(220px, 1fr));
  gap: var(--sp-xl);
  align-items: start;
}
.member-card {
  background: transparent;
  border: 0;
  border-radius: 0;
  overflow: visible;
  box-shadow: none;
  display: flex;
  flex-direction: column;
}
.member-card__photo {
  width: 100%;
  aspect-ratio: 4 / 5;
  overflow: hidden;
  background: transparent;
  flex-shrink: 0;
}
.member-card__photo img {
  width: 100%;
  height: 100%;
  object-fit: cover;
  object-position: center center;
  display: block;
}
.member-card__photo img[alt*="Ben Bellanger"] {
  object-position: 50% 50%;
}
.member-card__info {
  padding: var(--sp-md) var(--sp-lg) var(--sp-lg);
  display: flex;
  flex-direction: column;
  align-items: center;
  text-align: center;
  gap: 6px;
}
.member-card__name {
  font-size: 1.2rem;
  font-weight: 600;
  margin: 0;
  color: var(--text);
}
.member-card__role {
  font-size: .9rem;
  font-weight: 600;
  color: var(--navy);
  margin: 0;
}
.member-card__role-link {
  color: inherit;
  text-decoration: none;
}
.member-card__role-link:hover {
  color: var(--navy);
  text-decoration: underline;
  text-underline-offset: 0.16em;
}
.member-card__email {
  display: flex;
  align-items: center;
  justify-content: center;
  gap: 8px;
  font-size: .84rem;
  color: var(--muted);
  margin: 0 0 var(--sp-sm);
  flex-wrap: wrap;
}
.member-card__email-link {
  color: inherit;
  text-decoration: underline;
  text-underline-offset: 0.16em;
}
.member-card__email-link:hover {
  color: var(--navy);
}
.copy-btn {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  width: 26px;
  height: 26px;
  color: var(--navy);
  background: var(--surface);
  border: 1px solid var(--border);
  border-radius: 4px;
  cursor: pointer;
  transition: background .15s, color .15s;
  flex-shrink: 0;
  padding: 0;
}
.copy-btn svg {
  width: 14px;
  height: 14px;
  display: block;
}
.copy-btn:hover { background: var(--navy); color: var(--white); }
.copy-btn[data-copied="true"] {
  background: var(--navy);
  color: var(--white);
}
.member-card .pi-links {
  margin-top: var(--sp-sm);
}

/* ---- Open Positions list ---- */
.open-positions {
  list-style: none;
  margin-top: var(--sp-lg);
  display: flex;
  flex-direction: column;
  gap: var(--sp-md);
}
.open-positions li {
  font-size: .95rem;
  line-height: 1.7;
  padding-left: 1.2em;
  position: relative;
  color: var(--text-mid);
}
.open-positions li::before {
  content: '—';
  position: absolute;
  left: 0;
  color: #E9B43A;
  font-weight: 700;
}
.open-positions li strong {
  color: var(--text);
}

.join-photos-caption {
  margin: var(--sp-lg) 0 var(--sp-sm);
  font-size: .9rem;
  color: var(--muted);
  font-style: italic;
}

.join-photos {
  margin-top: 0;
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: var(--sp-md);
  align-items: start;
}
.join-photos img {
  width: 100%;
  display: block;
  border: 1px solid var(--border);
  box-shadow: var(--shadow-sm);
  object-fit: contain;
  background: var(--white);
}

/* ---- People ---- */
.pi-card {
  display: grid;
  grid-template-columns: 280px 1fr;
  gap: var(--sp-xl);
  background: var(--white);
  border: 1px solid var(--border);
  border-radius: var(--r-xl);
  padding: var(--sp-xl);
  box-shadow: var(--shadow-sm);
  margin-bottom: var(--sp-2xl);
}
.pi-card__photo {
  width: 100%; aspect-ratio: 1;
  border-radius: var(--r-lg);
  background: var(--surface);
  display: flex; align-items: center; justify-content: center;
  overflow: hidden; border: 1px solid var(--border);
}
.pi-card__initials {
  font-family: var(--font-display);
  font-size: 5rem;
  color: var(--navy);
  opacity: .14;
  user-select: none;
}
.pi-card h2 { font-size: 2.4rem; margin-bottom: 4px; letter-spacing: -.01em; }
.pi-card__role { color: var(--amber); font-weight: 500; margin-bottom: 4px; }
.pi-card__institution { font-size: .84rem; color: var(--muted); margin-bottom: var(--sp-lg); }
.pi-card__bio { font-size: .94rem; line-height: 1.8; margin-bottom: var(--sp-lg); }
.pi-card__bio p { max-width: none; margin-bottom: 14px; }
.pi-card__bio p:last-child { margin-bottom: 0; }
.pi-card__edu h5 { color: var(--muted); margin-bottom: 12px; }
.pi-card__edu ul { list-style: none; }
.pi-card__edu li {
  font-size: .9rem; color: var(--muted);
  padding: 7px 0;
  border-bottom: 1px solid var(--border);
  display: flex; gap: var(--sp-md); align-items: baseline;
}
.pi-card__edu li:last-child { border-bottom: none; }
.pi-card__edu li strong { color: var(--text); font-weight: 600; min-width: 110px; flex-shrink: 0; }

.pi-links { display: flex; gap: 10px; flex-wrap: wrap; margin-top: var(--sp-md); }
.pi-link {
  display: inline-flex;
  align-items: center;
  gap: 6px;
  font-size: .9rem;
  font-weight: 700;
  color: var(--navy);
  background: var(--surface);
  padding: 10px 16px;
  border-radius: var(--r-sm);
  border: 1px solid var(--border);
  box-shadow: var(--shadow-sm);
  transition: all .2s;
}
.pi-link:hover { background: var(--navy); color: var(--white); border-color: var(--navy); transform: translateY(-1px); }

.positions-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: var(--sp-md);
}
.position-card {
  background: var(--white);
  border: 1px solid var(--border);
  border-radius: var(--r-md);
  padding: var(--sp-lg);
  transition: box-shadow .22s, transform .22s;
}
.position-card:hover { box-shadow: var(--shadow-md); transform: translateY(-3px); }
.position-card__icon {
  width: 48px; height: 48px;
  border-radius: var(--r-sm);
  background: var(--amber-pale);
  display: flex; align-items: center; justify-content: center;
  margin-bottom: var(--sp-md);
  font-size: 1.4rem;
}
.position-card h4 { margin-bottom: 8px; }
.position-card p  { font-size: .9rem; color: var(--muted); line-height: 1.65; }
.position-card .btn { margin-top: var(--sp-md); font-size: .82rem; padding: 9px 18px; }

/* ---- News timeline ---- */
#news.section {
  padding-top: var(--sp-lg);
  padding-bottom: var(--sp-lg);
}
#news { scroll-margin-top: calc(var(--nav-h) + 12px); }

.news-timeline {
  position: relative;
  padding-left: 52px;
  max-width: 800px;
}
.news-timeline::before {
  content: '';
  position: absolute;
  left: 16px; top: 12px; bottom: 0;
  width: 1px;
  background: var(--border);
}
.tl-item { position: relative; margin-bottom: var(--sp-xl); }
.tl-item::before {
  content: '';
  position: absolute;
  left: -44px; top: 8px;
  width: 10px; height: 10px;
  border-radius: 50%;
  background: #E9B43A;
  border: 3px solid var(--off-white);
  box-shadow: 0 0 0 1.5px #E9B43A;
}
.tl-item__date {
  font-size: .72rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: .1em;
  color: var(--muted);
  margin-bottom: 8px;
  display: flex; align-items: center; gap: 10px;
}
.tl-item h4 { font-size: 1.05rem; margin-bottom: 8px; }
.tl-item p  { font-size: .92rem; color: var(--muted); line-height: 1.7; }
.tl-item a  { color: var(--navy); font-weight: 500; }
.tl-item a:hover { color: var(--amber); }

/* ---- Contact ---- */
.contact-grid {
  display: grid;
  grid-template-columns: 1fr 1.4fr;
  gap: var(--sp-xl);
  align-items: start;
}
.contact-block { margin-bottom: var(--sp-lg); }
.contact-block__label {
  font-size: .68rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: .12em;
  color: var(--muted);
  margin-bottom: 6px;
}
.contact-block__value { font-size: .96rem; line-height: 1.62; }

.form-card {
  background: var(--white);
  border: 1px solid var(--border);
  border-radius: var(--r-xl);
  padding: var(--sp-xl);
  box-shadow: var(--shadow-sm);
}
.form-card h3 { font-size: 1.8rem; margin-bottom: var(--sp-sm); letter-spacing: -.01em; }
.form-card > p { font-size: .87rem; color: var(--muted); margin-bottom: var(--sp-lg); max-width: none; }
.form-group { margin-bottom: var(--sp-md); }
.form-label { display: block; font-size: .83rem; font-weight: 500; margin-bottom: 6px; }
.form-control {
  width: 100%;
  padding: 12px 14px;
  border: 1px solid var(--border);
  border-radius: var(--r-sm);
  font-size: .94rem;
  font-family: var(--font-sans);
  background: var(--white);
  color: var(--text);
  transition: border-color .2s, box-shadow .2s;
  outline: none;
  -webkit-appearance: none;
}
.form-control:focus {
  border-color: var(--navy);
  box-shadow: 0 0 0 3px rgba(13,43,85,.1);
}
textarea.form-control { min-height: 120px; resize: vertical; }
select.form-control {
  background-image: url("data:image/svg+xml,%3Csvg xmlns='http://www.w3.org/2000/svg' width='12' height='8'%3E%3Cpath d='M1 1l5 5 5-5' stroke='%236B7280' stroke-width='1.5' fill='none' stroke-linecap='round'/%3E%3C/svg%3E");
  background-repeat: no-repeat;
  background-position: right 14px center;
  padding-right: 40px;
}

/* ---- Join cards ---- */
.join-grid {
  display: grid;
  grid-template-columns: repeat(3, 1fr);
  gap: var(--sp-md);
  margin-bottom: var(--sp-2xl);
}
.join-card {
  background: var(--white);
  border: 1px solid var(--border);
  border-radius: var(--r-md);
  padding: var(--sp-lg);
}
.join-card__num {
  font-family: var(--font-display);
  font-size: 3.2rem;
  color: var(--navy-mid);
  opacity: .4;
  line-height: 1;
  margin-bottom: 14px;
  font-weight: 300;
}
.join-card h4 { margin-bottom: 10px; font-size: 1.05rem; }
.join-card p  { font-size: .9rem; color: var(--muted); line-height: 1.7; }
.join-card ul { list-style: none; margin-top: var(--sp-sm); }
.join-card ul li {
  font-size: .87rem; color: var(--muted);
  padding: 5px 0;
  border-bottom: 1px solid var(--border);
  display: flex; gap: 8px;
}
.join-card ul li::before { content: '→'; color: var(--amber); font-weight: 700; }
.join-card ul li:last-child { border-bottom: none; }

/* ---- Footer ---- */
.site-footer {
  background: var(--dark);
  color: rgba(255,255,255,.42);
  padding: var(--sp-xl) var(--sp-lg);
}
.site-footer__grid {
  max-width: var(--max-w);
  margin: 0 auto;
  display: grid;
  grid-template-columns: 2fr 1fr 1fr 1fr;
  gap: var(--sp-xl);
  padding-bottom: var(--sp-xl);
  border-bottom: 1px solid rgba(255,255,255,.06);
}
.footer-brand__name {
  font-family: var(--font-display);
  font-size: 1.4rem;
  color: var(--white);
  margin-bottom: 6px;
}
.footer-brand p { font-size: .82rem; line-height: 1.72; }
.footer-brand a { color: rgba(255,255,255,.42); transition: color .2s; }
.footer-brand a:hover { color: var(--white); }
.footer-nav__title {
  font-size: .64rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: .16em;
  color: rgba(255,255,255,.22);
  margin-bottom: var(--sp-md);
}
.footer-nav ul { list-style: none; display: flex; flex-direction: column; gap: 10px; }
.footer-nav a { font-size: .84rem; color: rgba(255,255,255,.42); transition: color .2s; }
.footer-nav a:hover { color: var(--white); }
.site-footer__bottom {
  max-width: var(--max-w);
  margin: var(--sp-md) auto 0;
  display: flex;
  justify-content: space-between;
  align-items: center;
  font-size: .74rem;
  color: rgba(255,255,255,.2);
  flex-wrap: wrap;
  gap: var(--sp-sm);
}

/* ---- Scroll reveal ---- */
[data-reveal] {
  opacity: 0;
  transform: translateY(24px);
  transition: opacity .7s ease, transform .7s ease;
}
[data-reveal].is-visible { opacity: 1; transform: none; }

[data-reveal-stagger] > * {
  opacity: 0;
  transform: translateY(20px);
  transition: opacity .6s ease, transform .6s ease;
}
[data-reveal-stagger].is-visible > * { opacity: 1; transform: none; }
[data-reveal-stagger].is-visible > *:nth-child(1) { transition-delay: 0s; }
[data-reveal-stagger].is-visible > *:nth-child(2) { transition-delay: .1s; }
[data-reveal-stagger].is-visible > *:nth-child(3) { transition-delay: .2s; }
[data-reveal-stagger].is-visible > *:nth-child(4) { transition-delay: .3s; }

/* ---- Utilities ---- */
.text-center { text-align: center; }
.mt-sm  { margin-top: var(--sp-sm); }
.mt-md  { margin-top: var(--sp-md); }
.mt-lg  { margin-top: var(--sp-lg); }
.mt-xl  { margin-top: var(--sp-xl); }
.mb-md  { margin-bottom: var(--sp-md); }
.mb-lg  { margin-bottom: var(--sp-lg); }
.mb-xl  { margin-bottom: var(--sp-xl); }

/* ---- Responsive ---- */
@media (max-width: 1024px) {
  .approach-grid { grid-template-columns: repeat(2, 1fr); }
  .site-footer__grid { grid-template-columns: 1fr 1fr; gap: var(--sp-lg); }
  .fp { grid-template-columns: 1fr; }
  .fp__left {
    flex-direction: row; gap: var(--sp-md);
    justify-content: flex-start;
    border-right: none; border-bottom: 1px solid rgba(255,255,255,.07);
    padding: var(--sp-lg);
  }
}
@media (max-width: 900px) {
  .research-area { grid-template-columns: 1fr; }
  .research-area--flip { direction: ltr; }
  .pi-card { grid-template-columns: 1fr; }
  .pi-card__photo { max-width: 200px; }
  .positions-grid { grid-template-columns: 1fr; }
  .join-grid { grid-template-columns: 1fr; }
  .contact-grid { grid-template-columns: 1fr; }
  .news-list-item { grid-template-columns: 88px 100px 1fr; }
  .news-list-item__arrow { display: none; }

}
@media (max-width: 640px) {
  :root { --sp-lg: 32px; --sp-xl: 60px; --sp-2xl: 80px; }
  .approach-grid { grid-template-columns: 1fr; }
  .site-footer__grid { grid-template-columns: 1fr; gap: var(--sp-lg); }
  .pub-item { flex-direction: column; gap: 6px; }
  .research-list-item { grid-template-columns: 52px 1fr; }
  .research-list-item__arrow { display: none; }
  .news-list-item { grid-template-columns: 1fr; gap: 5px; padding: 16px var(--sp-sm); }
  .news-list-item__date { display: none; }
  .hero__headline { font-size: clamp(3rem, 13vw, 4.5rem); }

  /* Mobile nav — hamburger only below 640px */
  .site-nav__links {
    display: none;
    position: fixed; top: var(--nav-h); left: 0; right: 0;
    background: rgba(10,20,40,.97);
    flex-direction: column;
    padding: var(--sp-sm) var(--sp-lg);
    gap: 0;
    border-bottom: 1px solid rgba(255,255,255,.08);
  }
  .site-nav.scrolled .site-nav__links { background: rgba(255,255,255,.98); }
  .site-nav.scrolled .site-nav__links a { color: var(--text-mid) !important; border-bottom: none; font-weight: 400; }
  .site-nav__links.is-open { display: flex; }
  .site-nav__links li { border-bottom: 1px solid rgba(255,255,255,.06); }
  .site-nav.scrolled .site-nav__links li { border-bottom-color: var(--border); }
  .site-nav__links a { display: block; padding: 14px 0; color: rgba(255,255,255,.72) !important; border-bottom: none !important; }
  .site-nav__hamburger { display: block; }
}